In 1943, the Axis powers decide that a show of strength might bully neutral Turkey into joining them. Their target is 2,000 British soldiers marooned on the island of Keros in the Aegean Sea. Rescue by the Royal Navy is impossible because of massive radar-directed guns on the nearby island of Navarone. Time is short, because the Germans are expected to launch an assault on the British forces.
Efforts to destroy the guns by aerial bombing have proved fruitless. So that six destroyers can pick up the stranded men, Commodore Jensen (James Robertson Justice) of Allied Intelligence gathers a team of commandos to sail to Navarone and destroy the guns. Led by Major Roy Franklin (Anthony Quayle), they are Captain Keith Mallory (Gregory Peck), renowned mountaineer; Colonel Andrea Stavrou (Anthony Quinn), from the defeated Greek army; Franklin's best friend Corporal Miller (David Niven), a former University Chemistry teacher and explosives expert; Greek-American Spyros Pappadimos (James Darren), a native of Navarone; and "Butcher" Brown (Stanley Baker), an engineer and expert knife fighter.
Disguised as Greek fishermen on a decrepit boat, they sail across the Aegean Sea. They are intercepted by a German boat and boarded. They attack and kill all the Germans and sink the patrol boat. During the remainder of the voyage, Mallory confides to Franklin that Andrea has sworn to kill him after the war, because he was inadvertently responsible for the deaths of Andrea's wife and children.
In a violent storm, the ship is wrecked and they lose part of their equipment but still manage to land on the island. Led by Mallory, who was recruited for his climbing skills, they scale the "unclimbable" cliff. But Franklin is badly injured; the injury later becoming infected with gangrene. They find that the cliff is, in fact, guarded after all. Miller suggests that they leave Franklin to be "well cared for" by the enemy. Mallory, who assumes command of the mission, feels that Franklin would be forced to reveal their plans, so he orders two men to carry the injured man on a stretcher.
Franklin tries to commit suicide, but Mallory lies to him, saying that their mission has been "scrubbed" and that a major naval attack will be mounted on the side of the island opposite the gun emplacement. They rendezvous with local resistance fighters, Spyros's sister Maria (Irene Papas) and her friend Anna (Gia Scala), who was captured, tortured, and escaped. She was so traumatized that she cannot speak and will not allow even Maria to see her scars.
The mission is continually dogged by German soldiers, and the group is captured by Oberleutnant Muesel (Walter Gotell) in the town of Mandrakos, when they try to find a doctor for Franklin. Muesel and Hauptsturmführer Sessler (George Mikell) of the SS fail to persuade the saboteurs to tell them where Miller's explosives are. Andrea pretends to grovel and beg for mercy, which surprises the Germans, allowing the group to overpower their captors. They escape in German uniform but leave Franklin behind to receive medical attention.
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